#1 F5.A12 FIELD95.A13 F6-10.A12 F7.A12 F1.A12 F2-3.A12 F4.A12 *
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Revised Oct. 9, 1994
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abdominal sternites progressively rising posteriorly towards apices
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of elytra (fig. 5); elytra flat, posterior elytra not sloping ventrally;
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tibiae produced apically into long, curved tooth (fig. 6)
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Subfamily SCOLYTINAE *A*
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abdominal sternites not rising posterior (fig. 7); posterior elytra sloping
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ventrally; tibia with small lateral spines *B*
#B F9.A12 F7.A12 F6-10.A12 backto #1
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head completely concealed from above; pronotum granulate anteriorly and
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punctate posteriorly (fig. 9); third tarsal segment not bilobed (fig. 10)
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Subfamily IPINAE *C*
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head partly visible from above; pronotum uniformly sculptured; third
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segment of tarsi bilobed (fig. 8): Subfamily HYLESININAE *D*
#D F11-12.A12 backto #B
^8 3
anterior elytral margin granulate,
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ridged (fig. 11) *2B*
^8 45
anterior elytral margin smooth,
^9 45
not ridged (fig. 12) *17B*
#2B F13-14.A12 backto #D
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anterior elytral margin almost
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straight (fig. 13) *16B*
^10 40
anterior elytral margin curved on
^11 40
each side of scutellum (fig. 14) *3B*
#3B F15-16.A12 backto #2B
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club distinctly 3-segmented (fig. 15) *4B*
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club fused, without distinct segments (fig. 16) *6B*
#4B F17-18.A12 backto #3B
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club pectinate (fig. 17): Genus Phloeotribus *41B*
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club not pectinate (fig. 18) *5B*
#5B F19-20.A12 backto #4B
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anterior pronotum granulate (fig. 19):
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Genus Phloeophthorus *45B*
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anterior pronotum not granulate (fig. 20):
^11 38
Genus Phthorophloeus *43B*
#6B F21-22.A12 backto #3B
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eyes distinctly anteriorly emarginate
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(fig. 21): Genus Phloeosinus *83B*
^12 45
eyes not emarginate (fig. 22) *7B*
#7B F23-24.A12 backto #6B
^9 2
prothoracic coxae distinctly separated
^10 2
by broad prosternal
^11 2
process (fig. 23) *8B*
^13 40
prothoracic coxae almost contiguous
^14 40
(fig. 24) *14B*
#8B F25-26.A12 backto #7B
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elytra sloping downward; abdominal sternites progressively rising posterior
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to the elytral apices (fig. 25) *9B*
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elytra regularly rounded; abdominal sternites flat, not rising
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posterior to the elytral apices (fig. 26) *10B*
#9B F27-28.A12 backto #8B
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pronotum and elytra with filiform setae (fig. 27): Genus Hylesinus *69B*
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pronotum and elytra with squamiform setae (fig. 28): Genus Leperisinus *71B*
#10B F29-30.A12 backto #8B
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elytra with squamiform setae (fig. 29); funicle 7-segmented *11B*
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elytra with squamiform and filiform setae (fig. 30); funicle 5 to 6,
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seldom 7-segmented *12B*
#11B F31.A12 F32.A12 backto #10B
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pronotum and elytra with thick, round, squamiform setae; scutellum
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visible (fig. 31): Genus Pteleobius *73B*
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elytra with long, contiguous, squamiform setae; pronotum with fine,
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filiform setae (fig 32): Genus Chaetoptelius *79B*
#12B backto #10B
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elytral declivity on second interstria with spiniform or blunt granules:
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Genus Hylastinus *77B*
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elytral declivity on second interstria with dense, filiform setae,
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granules absent *13B*
#13B F33-34.A12 backto #12B
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setae on pronotum inclined medially (fig. 33); funicle 5-segmented:
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Genus Xylechinus *75B*
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setae on pronotum inclined concentrically (fig. 34); funicle 6-segmented:
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Genus Kissophagus *81B*
#14B F35-36.A12 backto #7B
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anterior pronotum margin not emarginate (fig. 35); funicle 6-segmented *15B*
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anterior pronotum margin medially emarginate (fig. 36); funicle 5-segmented:
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Genus Dendroctonus *59B*
#15B F37-38.A12 backto #14B
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interstriae with dense, filiform setae (fig. 37); club nearly conical:
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Genus Hylurgus *57B*
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interstriae with rows of sparsely distributed filiform setae (fig. 38);
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club oval: Genus Tomicus (Blastophagus) *55B*
#16B F39-40.A12 backto #2B
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eyes bipartite (fig. 39); club without sutures: Genus Polygraphus *65B*
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eyes medially emarginate anteriorly (fig. 40); club with sutures:
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Genus Carphoborus *61B*
#17B F41-42.A12 backto #D
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pronotum wider than long, anteriorly constricted; elytral posteriorly
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broadened (fig. 41): Genus Hylurgops *53B*
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pronotum as wide as long; elytra not broadened posteriorly (fig. 42):
^19 1
Genus Hylastes *47B*
#41B F99.A12 F100-101.A12 backto #4B
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Phloeotribus: 1.8 to 2.4 mm; body black/brown; antennae inserted close together;
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club 3-segmented, laterally pectinate, with dense, filiform setae (fig. 99);
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elytra with fine, grey,
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squamiform, and short, filiform
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setae; elytra with transverse, medial
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band of brown, squamiform setae; pronotum with long,
^16 1
filiform setae (fig. 100); 2 to 2.5 mm:
^17 1
Phloeotribus scarabaeoides BERNARD *
^28 1
elytra with dense, fine, grey, filiform setae; interstriae with rows of
^29 1
light-brown, filiform setae (fig. 101); filiform setae on pronotum not
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longer than setae on elytra; 1.8 to 2 mm: Phloeotribus caucasicus REITTER *
#45B backto #5B
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Phloeophthorus: 1.5 to 1.8 mm; body dark-grey, with short, grey, filiform
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setae; antennae and legs light brown; frons flat with 2 tubercles; semi-
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circular depression above the mandibles; pronotum wider than long, with sparse,
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large puncture; pronotum anterior with 3 or 4 small, bilaterally distributed
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tubercles, sometimes with short, medial, longitudinal ridge; elytra 1.5 x as
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long as wide, with rough punctures; striae with short, blond, filiform setae
^7 1
lying flat on elytra; interstriae with rows of long, grey, filiform setae.
^9 1
anterior elytral margin granulate, submarginal tubercles absent; pronotum
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with light brown, filiform setae and fine granules; elytra 2 x as long as wide;
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body black; 1.5 to 1.8 mm: Phloeophthorus rhododactylus MARSHAM *
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anterior elytral margin granulate with submarginal row of small tubercles *45C*
#45C F103.A12 backto #45B
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elytra without rows of punctures (fig. 103); frons of male with medial groove;
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female's rounded, roughly punctured, dull; body black with fine, light brown,
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filiform setae; 1.5 to 1.8 mm: Phloeophthorus brevicollis KOLENATI *
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elytra with rows of rough punctures and find, white setae; pronotum densely
^26 1
punctured, dull, with long, filiform setae; male frons with smooth, medial
^27 1
groove and 2 small tubercles; 1.5 mm: Phloeophthorus sharpi GUILLEBEAU *
#43B F102.A12 backto #5B
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Phthorophloeus: club distinctly 3-segmented, slightly laterally pectinate
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pronotum without tubercles; elytra with rows of rough punctures; interstriae
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on declivity with rows of small setae;
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posterior pronotum narrower than anterior elytra (fig. 102); elytra with
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rows of rough punctures; interstriae with blond setae; frons of male concave,
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female's round; body black, head and pronotum dark brown, antennae and
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tarsi reddish brown; 1.8 to 2.2 mm (Picea): Phthorophloeus spinulosus REY *
#83B backto #6B
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Phloeosinus: 1.2 to 4 mm; eyes medial, laterally emarginate; funicle
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segmented; club long, oval, with 2 lateral indentations (fig. 154); anterior
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pronotum constricted; anterior elytral margin emarginate, with granules;
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interstriae broad and wrinkled; 1st, 3rd, 5th, and 7th interstriae on
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declivity with rows of pointed tubercles or teeth; interstriae 1, 3, 5, and 7
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on female declivity with rows of small tubercles; other sexual dimorphism:
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topical morphology of frons; monogamous; host: Cypressus spp.
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Male declivity with 4 broad spines (fig. 155); female declivity with 2 rows
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of 4 conical spines (fig. 156); pronotum with partial, medial, longitudinal
^11 1
ridge without punctures; male frons with medial groove; body reddish brown;
^12 1
antennae and legs light brown; head black; 4 to 4.5 mm.
^13 1
Phloeosinus armatus REITTER *
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spines on elytral declivity absent, with only 1 to 2 rows of granules *83C*
#83C backto #83B
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3rd interstria not ridged on declivity; sexual dimorphism: male with row of
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find, dense tubercles on 1st and 3rd interstriae on declivity (fig. 157);
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body brown to black; antennae, elytra, and legs reddish brown; 2 to 2.5 mm:
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Phloeosinus aubei PERRIS *
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3rd interstria ridged on elytral declivity *83D*
#83D backto #83C
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2nd interstria narrower than others; body light brown; antennae and tarsi
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reddish brown; 1.5 to 2.4 mm: Phloeosinus thujae PERRIS *
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2nd interstria as wide as others *83E*
#83E backto #83D
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3rd interstria on declivity with row of small granules, larger in male;
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male frons flat, with fine, medial, longitudinal ridge; female frons round;
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body black; 2 mm: Phloeosinus stoechleini SCHEDL *
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1st, 2nd, and 3rd interstriae on declivity with large tubercles (fig. 158);
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body black to brown, elytra reddish brown; antennae and tarsi reddish brown;
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2.5 to 3.5 mm: Phloeosinus rudis *
#69B backto #9B
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Hylesinus: 2.2 to 6 mm; funicle 7-segmented; club long, pointed, 3-sutured
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(fig. 140); lateral elytral margin with furrow; abdominal sternites
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progressively rising posteriorly; body black; antennae and tarsi reddish brown.
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anterior pronotum with tubercles (fig. 141); pronotum and elytra with long
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filiform setae; body black; antennae and tarsi red; 2.5 to 3.2 mm:
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Hylesinus oleiperda FABRICIUS *
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anterior pronotum without tubercles (fig. 142); pronotum and elytra with short,
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thick filiform setae; elytra with indented striae; funicle with long, filiform
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setae; body black; antennae and tarsi reddish brown; 3.8 to 6 mm:
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Hylesinus crenatus FABRICIUS *
#71B bacto #9B
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Leperisinus: 2.5 to 3.5 mm; posterior margin of 1st and 2nd sternite straight,
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2nd sternite smaller than 3rd and 4th sternite together; funicle 7-segmented;
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club long, pointed, 2-sutured (fig. 143); monogamous; host: deciduous trees.
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anterior pronotum with granules; pronotum and elytra with light brown, over-
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lapping, squamiform setae (fig. 144); anterior elytra on 2nd interstria with
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tubercles; other interstriae with rows of granules; sexual dimorphism: males
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stridulate; shape of terminal abdominal segment as in fig. 145 and 146;
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2.5 to 2.5 mm: Leperisinus varius FABRICIUS *
#73B bacto #11B
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Pteleobius: 1.8 to 2.2 mm; body cylindrical; abdominal segments not rising
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posteriorly; posterior elytra sloping downward; tibiae furrowless; elytra and
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pronotum with short, round, squamiform setae; scutellum visible; monogamous;
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host: elm; transfer spores of Ceratocystis ulmi.
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interstriae with short, thick, filiform setae, longer than squamiform setae
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(fig. 147); elytra with dense, light brown to white, and dark brown, squamiform
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setae; 2nd interstria not reaching posterior elytral margin; club brown:
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Pteleobius kraatzi EICHHOFF *
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interstriae with small, filiform setae (fig. 148); squamiform setae fine,
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light brown; elytra with dark brown, medial W-shaped band of squamiform setae;
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2nd interstria reaches posterior elytral margin; club black with 3 medial
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bowed sutures (fig. 149): Pteleobius vittatus FABRICIUS *
#79B backto #11B
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Chaetoptelius: 2.4 to 3.5 mm; elytra with brown or yellow-grey squamiform
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setae; dark brown squamiform setae distributed with bilateral symmetry;
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sexual dimorphism: female frons round with sparse filiform setae and fine
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medial, longitudinal ridge; male frons concave with dense filiform setae;
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monogamous.
^7 1
elytra with sparse, long, dark brown filiform setae and find squamiform setae,
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darker on declivity (fig. 152); ventral body with dense, white, filiform setae;
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body reddish brown to dark brown; antennae and legs light brown; 2.4 to 3.5 mm:
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Chaetoptelius vestitus REY *
#77B bacto #12B
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Hylastinus: 2 to 2.5 mm; funicle 7-segmented; club oval, 2-sutured (fig. 151A);
^2 1
pronotum irregularly punctured; elytra with rough striae; interstriae with rows
^3 1
of filiform setae; ventral mesothorax with dense, light brown filiform setae;
^4 1
monogamous.
^6 1
elytra with dense, fine, filiform setae; pronotum roughly punctured, dull,
^7 1
with medial, longitudinal, ridged band (fig. 151B); body brown; elytra,
^8 1
antennae, and legs reddish brown; 2 to 2.5 mm:  Hylastinus obscurus MARSHAM *
^10 1
elytra without filiform setae; interstriae with 3 rows of filiform setae;
^11 1
pronotum roughly punctured, without medial, longitudinal ridged band; body
^12 1
black to brown; elytra, antennae and legs reddish brown; 2 to 2.5 mm:
^13 1
Hylastinus tiliae SEMENOV *
#75B bacto #13B
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Xylechinus: funicle 5-segmented; club with 3 bowed sutures (fig. 150); pronotum
^2 1
with flat lying filiform setae; elytra anteriorly constricted before anterior
^3 1
margin; filiform setae of 1st interstria lighter and more densely distributed
^4 1
than other interstriae; monogamous; host: conifers.
^6 1
pronotum narrower than elytra, anterior slightly constricted, with fine
^7 1
punctures and filiform setae (fig. 151); elytra with fine, squamiform setae;
^8 1
body dark brown; antennae and legs reddish brown; 2.2 to 2.5 mm:
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Xylechinus pilosus RATZEBURG *
#81B backto #13B
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Kissophagus: anterior pronotum with fine granules; pronotum with concentric,
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filiform setae; elytra with thick, squamiform setae; interstriae with row of
^3 1
light brown, short, thick, filiform setae; body reddish brown; antennae and
^4 1
legs light brown; monogamous; host: ivy.
^6 1
anterior pronotum margin smooth; pronotum with sparse, fine granules; elytra
^7 1
with fine rows of punctures; interstriae with fine punctures, wrinkled, with
^8 1
row of short, thick, filiform setae; 2 to 2.4 mm:
^9 1
Kissophagus hederae SCHMIDT *
^11 1
anterior pronotal margin ridged, with tubercles; pronotum granulate (fig. 153);
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elytral striae rough; interstriae with rough punctures, wrinkled, and with row
^13 1
of thick, filiform setae; 2 to 2.3 mm: Kissophagus novaki REITTER *
#59B F127.A12 F128.A12 backto #14B
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Dendroctonus: funicle 5-segmented; club 4-segmented, nearly square, flat
^2 1
(fig. 127); anterior pronotum and elytral margin emarginate; anterior elytral
^3 1
margin granulate; punctures of elytral striae large, flat; interstriae with
^4 1
irregularly distributed granules; monogamous; host: conifers.
^6 1
anterior pronotum constricted (fig. 128); pronotum with irregularly distributed
^7 1
punctures; body black, shiny, with long, light brown, filiform setae; antennae
^8 1
and tarsi brown; male 5.5 to 6.5 mm; female 7 to 9 mm:
^9 1
Dendroctonus micans KUGELANN *
#57B F124.A12 backto #15B
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Hylurgus: 2 to 5.7 mm; body cylindrical, black to brown; antennae and legs
^2 1
light brown; funicle 6-segmented; club conical (fig. 124); metathorax as long
^3 1
as abdomen; anterior elytral margin granulate, not ridged; pronotum longer
^4 1
than wide; pronotum and elytra with long, fine, filiform setae; interstriae
^5 1
with dense, filiform setae; sexual dimorphism: male 2nd interstria with
^6 1
granules; monogamous; host: pines.
^8 1
body with dense, filiform setae; medial, longitudinal ridge on pronotum
^9 1
absent *57C*
^11 1
body with long, sparse, filiform setae; pronotum with medial, longitudinal
^12 1
ridge without punctures; elytra 2 x as long as pronotum; body brown to black,
^13 1
shiny; 2 mm: Hylurgus longulus KOLENATI *
#57C F125-126.A12 F125-126.A12 backto #57B
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filiform setae longer on pronotum than on elytra (fig. 125); pronotum
^2 1
punctured; punctures of striae rough, regularly distributed; body black to
^3 1
brown; antennae and tarsi reddish; 4 to 5.7 mm: Hylurgus ligniperda FABRICIUS *
^5 1
filiform setae on pronotum as long as on elytra (fig. 126); pronotum punctured;
^6 1
punctures of striae regular, fine; body dark brown; 3.5 to 3.8 mm:
^7 1
Hylurgus micklitzi WACHTL *
#55B F121.A12 F122-123.A12 backto #15B
^1 1
Tomicus (Blastophagus): 3.4 to 4.8 mm; funicle 6-segmented; club elongate,
^2 1
oval (fig. 121); posterior elytra before declivity slightly wider; elytra
^3 1
shiny, with sparse, filiform setae; pronotum wider than long; anterior elytral
^4 1
margin with granules; monogamous; hosts: pines.
^6 1
2nd interstria with row of granules on elytral declivity (fig. 122); frons
^7 1
sparsely punctured; sexual dimorphism: males stridulate; body black; antennae,
^8 1
elytra, and legs reddish-brown; 3.5 to 4 mm: Tomicus minor HARTIG *
^10 1
granules of 2nd interstria on elytral declivity absent (fig. 123);
^11 1
frons densely punctured; sexual dimorphism: males stridulate;
^12 1
body black; antennae, elytra and legs reddish-brown; 3.5 to
^13 1
4.8 mm: Tomicus piniperda L. *
#65B backto #16B
^1 1
Polygraphus: 1.2 to 3.5 mm; body cylindrical; eyes bipartite; funicle 5 or
^2 1
6 segmented; club flat, without sutures; 3rd tarsal segment cylindrical;
^3 1
anterior elytral margin with small tubercles; 1st sternite as long as terminal
^4 1
sternite; 2nd, 3rd, and 4th sternite small; sexual dimorphism: male frons
^5 1
rounded, with short, filiform setae and 2 medial tubercles (fig. 134); female
^6 1
frons with corona of short, filiform setae (fig. 135); polygamous; on conifers.
^8 1
funicle 6-segmented; club large oval (fig. 136); legs brown to black; antennae
^9 1
and tarsi light brown; pronotum densely punctured, dull, with ridged, medial,
^10 1
longitudinal band; granules on interstriae absent; 2.5 to 3 mm:
^11 1
Polygraphus grandiclava THOMSON *
^13 1
funicle 5-segmented; interstriae with rows of fine granules; antennae and
^14 1
legs light brown, pronotum shiny *65C*
#65C backto #65B
^1 1
club distally pointed; funicle 5-segmented (fig. 137); anterior pronotal margin
^2 1
emarginate (fig. 138); pronotum fine punctured, with short, medial, longi-
^3 1
tudinal band without punctures; body brown, with dense, short, light brown,
^4 1
squamiform setae; head black; 2.2 to 3 mm: Polygraphus poligraphus L. *
^6 1
club distally more rounded than pointed (fig. 139) *67B*
#67B backto #65C
^1 1
1.2 to 2.2 mm; frons roughly punctured; pronotum dense, with rough punctures
^2 1
and fine, medial, longitudinal ridge (fig. 139A); anterior pronotal margin
^3 1
straight; elytra with fine striae; declivity reddish brown, with shiny,
^4 1
squamiform setae: Polygraphus subopacus *
^6 1
2.3 to 3 mm; frons roughly punctured, dull; pronotum dense, rough punctate,
^7 1
dull, with narrow, medial, longitudinal ridge; elytra with striae; body black
^8 1
to brown, dull, with long, light brown, squamiform setae:
^9 1
Polygraphus punctifrons THOMSON *
#61B F129.A12 F130.A12 backto #16B
^1 1
Carphoborus: 1.2 to 2.1 mm; body black, dull, with dense, flat-lying, grey,
^2 1
squamiform setae; tarsi light brown; eyes laterally emarginate; funicle 5-
^3 1
segmented; club 4-sutured (fig. 129); anterior elytral margin granulate;
^4 1
sexual dimorphism: male frons with 2 small, medial tubercles, and dense, short,
^5 1
filiform setae; female frons punctured and with corona of short, filiform
^6 1
setae; polygamous; host: conifers.
^8 1
anterior pronotum not constricted
^9 1
(fig. 130); anterior pronotal margin
^10 1
with dense, long, filiform setae;
^11 1
club oval; body black, with brown,
^12 1
squamiform setae; antennae and legs
^13 1
light red; ventral body with fine,
^14 1
dense, white, filiform setae;
^15 1
1.8 to 2 mm:
^16 1
Carphoborus perrisi CHAPUIS *
^23 1
anterior pronotum constricted *61C*
#61C F131.A12 backto #61B
^1 1
3rd interstria on elytral declivity not ridged; 1st and 7th interstriae on
^2 1
declivity contiguous, ridged, and with pointed tubercles (fig. 131); 3rd and
^3 1
5th interstriae do not reach posterior elytral margin; body black; elytra
^4 1
dark brown with grey, squamiform setae and rough striae; antennae and tarsi
^5 1
reddish brown; 1.4 to 1.6 mm: Carphoborus pini EICHHOFF *
^9 1
3rd interstria ridged on elytral declivity *61D*
#61D backto #61C
^1 1
3rd interstria roughly ridged on elytral declivity; declivity not rounded *63B*
^3 1
3rd interstria sightly ridged on elytral declivity; declivity rounded *63C*
#63B F132-133.A12 backto #61D
^18 1
1st, 3rd, 5th, 7th, and 9th interstriae ridged on elytral declivity; 1st, 3rd
^19 1
and 9th interstriae posteriorly contiguous; body black; elytra and legs
^20 1
reddish brown; 1.7 to 1.9 mm: Carphoborus rossicus SEMENOV *
^22 1
1st, 3rd and 9th interstriae slightly ridged on elytral declivity (fig. 132);
^23 1
5th and 7th interstriae posteriorly contiguous; body black to brown; elytra
^24 1
with light brown, squamiform setae; 1.4 to 2 mm:
^25 1
Carphoborus cholodkovskyi SPESSIVTSEFF *
#63C F132-133.A12 backto #61D
^15 1
1st, 3rd and 9th interstriae posteriorly contiguous, ridged on elytral
^16 1
declivity, granules absent (fig. 133); other interstriae not reaching post-
^17 1
erior elytral margin; body black, with fine, yellow or grey, squamiform setae;
^18 1
antennae and tarsi light brown; 1.2 to 1.8 mm: Carphoborus minimus FABRICIUS *
^12 1
3rd and 9th interstriae posteriorly contiguous; other interstriae not
^13 1
reaching posterior elytral margin *63D*
#63D backto #63C
^1 1
1st, 3rd, 5th, 7th and 9th interstriae ridged on elytral declivity, and with
^2 1
small granules; body shiny, black to brown, with grey, squamiform setae;
^3 1
1.8 to 2.1 mm: Carphoborus teplouchovi SPESSIVTSEFF *
^5 1
1st, 3rd and 9th interstriae ridged on elytral declivity; granules absent;
^6 1
8th interstria posteriorly constricted; elytra with rough striae; antennae
^7 1
light brown, club dark brown; 1.6 to 1.8 mm: Carphoborus marani PFEFFER *
#53B F118.A12 F119-120.A12 backto #17B
^1 1
Hylurgops: 2.5 to 5.5 mm; head slightly elongate; funicle 7-segmented (fig.
^2 1
118); pronotum wider than long, anteriorly constricted, often with narrow,
^3 1
medial, longitudinal ridge; monogamous; host: conifers.
^5 1
interstriae as wide as striae, with short, filiform setae, and rows
^6 1
of granules (fig. 119); pronotum slightly narrower than elytra;
^7 1
lateral pronotum roughly punctured; body black; pronotum,
^8 1
elytra, and legs reddish brown; 2.5 to 3.2 mm:
^9 1
Hylurgops palliatus GYLLENHAL *
^12 1
interstriae at similar level, but wider than striae;
^13 1
declivity with rows of granules and filiform setae
^14 1
(fig. 120); pronotum narrower than elytra,
^15 1
densely punctured; with medial, longitudinal band
^16 1
without punctures; body dark brown; antennae and
^17 1
tarsi reddish brown; 4.5 to 5 mm:
^18 1
Hylurgops glabratus ZETTERSTEDT *
#47B F104.A12 backto #17B
^1 1
Hylastes: 2 to 5 mm; body dark reddish-brown to black; head elongate; frons
^2 1
sometimes with medial, longitudinal ridge; funicle 7-segmented, club oval,
^3 1
3-sutured (fig. 104); posterior margin of pronotum straight; anterior 2/3-3/4
^4 1
elytra parallel; monogamous; host: conifers.
^6 21
body longer than 3 mm *47C*
^8 21
body 3 mm or smaller *49B*
#47C F105-106.A12 backto #47B
^1 1
elytra more than 2 x as long as wide *47D*
^16 41
elytra at most 2 x as long as wide
^17 41
(fig. 105); pronotum as long as wide,
^18 41
anteriorly constricted; elytra dull,
^19 41
roughly punctured; interstriae wrinkled,
^20 41
with rows of filiform setae; elytral
^21 41
declivity with rows of white, squamiform
^22 41
and filiform setae; frons with
^23 41
transverse furrow at posterior origin
^24 41
of medial, longitudinal ridge
^25 41
(fig. 106); body black, pronotum reddish
^26 41
brown; 3.2 to 4.8 mm:
^27 41
Hylastes cunicularius ERICHSON *
#47D F108.A12 F107-110.A12 F111-112.A12 backto #47C
^18 1
pronotum longer than wide, narrower than elytra; pronotum punctate, except for
^19 1
partial, medial, longitudinal smooth band (fig. 107); frons with medial
^20 1
longitudinal ridge (fig. 108); elytral striae rough; interstriae with short,
^21 1
light brown, filiform setae; elytra black, dull; antennae and tarsi reddish
^22 1
brown; sexual dimorphism: male terminal tergite with medial depression (figs.
^23 1
109, 110); 4.5 to 4.8 mm: Hylastes ater PAYKUL *
^26 1
pronotum as long as wide, as wide as elytra (fig. 111); frons with dense, rough
^27 1
punctures (fig. 112); short filiform setae on interstriae; body black; tarsi
^28 1
brown; antennae reddish brown; 4.5 to 5.2 mm:
^29 1
Hylastes batnensis BRISOUR DE BARNEVILLE *
#49B backto #47B
^1 1
interstriae narrower than striae, with setae *49C*
^3 1
interstriae not narrower than striae *51B*
#49C F115.A12 F113-114.A12 backto #49B
^1 1
pronotum as long as wide, with long, blond, filiform setae (fig. 113); frons
^2 1
with short, medial, longitudinal ridge (fig. 114); elytra with rough striae
^3 1
and short, filiform setae; interstriae with row of granules and filiform setae;
^4 1
body dark brown; elytra, antennae and legs reddish brown; 2 to 3 mm:
^5 1
Hylastes attenuatus ERICHSON *
^6 1
Pronotum longer than wide, posteriorly constricted, with medial, longitudinal
^9 1
band without punctures (fig. 115); elytra with striae; interstriae with rows
^10 1
of squamiform setae; elytral suture slightly depressed; frons with indistinct,
^11 1
medial ridge; body brown to black; antennae and legs reddish brown; 3 mm:
^12 1
Hylastes linearis ERICHSON *
#51B F116-117.A12 backto #49B
^1 1
frons without longitudinal ridge (fig. 116); pronotum as long as wide,
^2 1
anteriorly constricted (fig. 117); elytra with flat striae; elytral declivity
^3 1
with filiform setae; body black; antennae and legs reddish brown; elytra
^4 1
dull; 2.5 to 3 mm: Hylastes opacus ERICHSON *
^6 1
frons with short, medial, longitudinal ridge *51C*
#51C backto #51B
^1 1
first 4 interstriae with at least 2 rows of setae; pronotum longer than wide,
^2 1
anteriorly constricted, with medial, longitudinal band without punctures; body
^3 1
black; elytra dark brown; 2.5 to 3 mm (south and middle Europe):
^4 1
Hylastes angustatus HERBST *
^6 1
anterior 1/4 of 1st interstria and all interstriae on declivity with 2 rows of
^7 1
setae; reticular interstriae with small granules; pronotum longer than wide,
^8 1
anteriorly and posteriorly constricted, with long, shiny, medial, longitudinal
^9 1
band without punctures; body brown; pronotum and elytra light brown; 2.6 mm:
^10 1
Hylastes scandinavicus LEKANDER *
#A F70-71.A12 F69.A12 backto #1
^1 1
Scolytus: 1.5 to 7 mm; eyes anterior slightly emarginate; club large, oval,
^2 1
sutures medially bowed in acute angle (fig. 69); pronotum large, anteriorly
^3 1
constricted, posterior and lateral margins sharply bordered; posterior elytra
^4 1
flat; abdomen rising posterior from 2nd sternite to apices; tibiae smooth
^5 1
laterally, produced distally into long, curved tooth; 3rd tarsal segments
^6 1
bilobed; sexual dimorphism: shape of head and setae, abdomen process;
^7 1
monogamous; host: deciduous trees; transfer spores of Ceratocystis ulmi.
^9 1
2nd abdominal sternite with protuberant process (fig. 70) *33C*
^11 1
2nd abdominal sternite without protuberant process (fig. 71) *35C*
#33C F72.A12 backto #A
^18 1
process of 2nd abdominal sternite small, inclined posteriorly (fig. 72);
^19 1
head: male's flat, with dense, long, filiform setae; female's arched,
^20 1
without setae; body dark brown to black; 2.5 mm: Scolytus kirschi SKALITZKY *
^23 5
process of 2nd abdominal sternite long, sturdy *33D*
#33D F73-74.A12 F75-76.A12 back to #33C
^1 1
posterior margin of 2nd, 3rd, and 4th sternite with small, lateral teeth (figs.
^2 54
73, 74); head: male's flat,
^3 58
with crown of long,
^4 57
filiform setae (fig. 74)
^5 58
female's arched, and
^6 58
nearly bald (fig. 73);
^7 58
body black; elytra
^8 54
reddish brown; 2 to 3.5 mm:
^9 49
Scolytus multistriatus MARSHAM *
^16 1
posterior margin of 2nd, 3rd and 4th sternite without lateral teeth (fig. 75);
^17 52
process of 2nd sternite long,
^18 56
distally slightly hooked;
^19 57
4th sternite with small,
^20 58
medial process; head of
^21 59
male flat, with dense,
^22 56
filiform setae (fig. 76);
^23 57
2 to 3 mm:
^24 54
Scolytus ensifer EICHHOFF *
#35C F77-78.A12 back to #A
^5 41
two different kinds of punctures on
^6 41
elytra; interstriae with sparsely
^7 41
distributed punctures (fig. 77) *35D*
^20 41
only one kind of puncture on elytra;
^21 41
striae dense (fig. 78) *37E*
#35D F79-81.A12 back to #35C
^1 36
frons with medial, longitudinal ridge,
^2 36
long in female (fig. 79), short in
^3 36
male (fig. 80); posterior part of 3rd
^4 36
sternite with broad, bilobate tubercle
^5 36
(fig. 81); body black, shiny;
^6 36
reddish brown; 5 to 6 mm:
^7 36
Scolytus ratzeburgi JANSON *
^27 41
frons without longitudinal ridge *35E*
#35E F82.A12 backto #35D
^18 1
lateral anterior pronotum with dense, rough punctures; punctures of rows and
^19 1
interstriae similar;; male frons flat, with long, blond, filiform setae;
^20 1
female frons round, with short, sparsely distributed, filiform setae (fig. 82);
^21 1
body dark brown to black; elytra with medial, transverse, dark band;
^22 1
3 to 3.2 mm:  Scolytus carpini RATZEBURG *
^25 1
lateral pronotum with fine punctures *35F*
#35F F83.A12 F84.A12 F85-86.A12 back to #35E
^1 1
interstriae irregularly punctate (figs. 83, 84); frons with dense, short,
^2 1
filiform setae; posterior margin of male terminal sternite with medial row
^3 1
of dense, long, filiform setae (fig. 85), absent in female (fig. 86); body
^4 1
black, elytra reddish brown, shiny; 3 to 6 mm: Scolytus scolytus FABRICIUS *
^6 1
interstriae with row of fine, regularly distributed punctures *37C*
#37C F87-88.A12 back to #35F
^1 1
rows of punctures on elytra dense (fig. 87) *37D*
^3 1
rows of punctures on elytra not dense (fig. 88); anterior pronotum fine,
^4 1
sparsely punctate, anteriorly constricted; almost bald; terminal abdominal
^5 1
sternite with medial, transverse depression in male, small, longitudinal
^6 1
depression in female; body black; elytra dark brown; 3 to 4 mm:
^7 1
Scolytus mali BECHSTEIN *
#37D F91-92.A12 F89-90.A12 back to #37C
^4 1
anterior elytra adjacent suture deeply depressed (fig. 89); frons with short
^5 1
furrow; male flat, with dense, filiform setae; female round, almost bald;
^6 1
body black; elytra reddish brown; 3.5 to 4.5 mm: Scolytus laevis CHAPUIS *
^8 1
elytra beside the suture not depressed (fig. 90); male frons flat, with long,
^9 1
filiform setae; female frons round, almost bald (figs. 91, 92); body black;
^10 1
elytra reddish brown; 2 to 3 mm: Scolytus pygmaeus FABRICIUS *
#37E backto #35C
^1 1
pronotum with large, oval, often fused punctures *39C*
^3 1
pronotum with small, round, dense, never fused punctures *39D*
#39C F95.A12 F93-94.A12 backto #37E
^1 1
male frons flat, with dense short, filiform setae (fig. 93); female frons
^2 1
round with sparsely distributed filiform setae; abdominal sternites
^3 1
progressively rising posteriorly (fig. 94); body black; elytra reddish brown
^4 1
with dark spots and dark medial band; 1.8 to 3 mm: Scolytus rugulosus *
^27 1
male frons round, with short, filiform setae; abdominal sternites constantly
^28 1
rising posteriorly (fig. 95); 5th abdominal sternite of male with dense,
^29 1
filiform setae; body black; elytra and anterior pronotum reddish brown; male
^30 1
3.2 mm, female 3.5 mm: Scolytus morawitzi SEMENOV *
#39D F96.A12 F97-98.A12 backto #37E
^26 1
abdomen sternites perpendicularly rising posteriorly (fig. 96); elytra with
^27 1
densely distributed rows of large punctures; male frons flat, with dense, long,
^28 1
filiform setae; female frons round with short, filiform setae; body black;
^29 1
elytra dark brown; antennae and legs reddish brown; 3 to 4.5 mm:
^30 1
Scolytus koenigi SCHEVYREV *
^1 1
abdominal sternites progressively rising posteriorly (fig. 97); elytra with
^2 1
rows of small punctures; male frons flat with short blond filiform setae with
^3 1
lateral brushes of long, blond, filiform setae above the mandibles; female
^4 1
frons round with sparsely distributed filiform setae (fig. 98); body black;
^5 1
elytra reddish brown; antennae and legs light brown; 2.5 to 3.5 mm:
^6 1
Scolytus intricatus RATZEBURG *
#C F43-44.A12 back to #B
^1 1
anterior pronotum with granules or spicules (fig. 43) *25B*
^3 1
pronotum entirely punctate, without granules or spicules (fig. 44) *27C*
#25B F45-46.A12 backto #C
^4 20
pronotum steeply arched (fig. 45) *25C*
^19 15
pronotum regularly rounded (fig. 46) *27D*
#25C F47-48.A12 backto #25B
^11 2
funicle 4-segmented (fig. 47) *25D*
^9 40
funicle 5-segmented (fig. 48) *27A*
#25D F49-50.A12 backto #25C
^1 1
anterior pronotal margin without protruding asperities (fig. 49) *25E*
^3 1
medial anterior pronotal margin with protruding asperities (fig. 50) *25F*
#25E back to #25D
^1 1
interstriae with rows of filiform or small squamiform setae; funicle
^2 1
4-segmented; club 3-sutured: Genus Cryphalus *105A*
^4 1
interstriae with rows of thick, squamiform setae; male funicle 3 to 4-
^5 1
segmented; club with 3 laterally indented sutures: Genus Hypothenemus *113A*
#25F backto #25D
^1 1
pronotum with asperities, broad anteriorly; tubercles irregularly distributed;
^2 1
club with bowed sutures: Genus Ernoporus *109A*
^4 1
pronotum with asperities, narrow anterior; spicules in 4 regular rows; club
^5 1
with straight sutures: Genus Cryphalops *111A*
#27A F51-52.A12 backto #25C
^1 1
club long, conical (fig. 51); elytral declivity adjacent suture with furrow;
^2 1
3rd or 4th interstria with granules: Genus Trypophloeus *99A*
^9 40
club broad (fig. 52) *27B*
#27B F53-54.A12 backto #27A
^1 1
prothoracic coxae separated by broad prosternal process (fig. 53); pronotum
^2 1
round, not longer than wide; anterior pronotal margin granulate:
^3 1
Genus Xylosandrus *159A*
^5 1
prothoracic coxae almost contiguous (fig. 54); anterior pronotal margin smooth:
^6 1
Genus Xyleborus *153A*
#27C F55.A12 backto #C
^1 1
funicle 2-segmented (fig. 55); sutures of antennal club indistinct:
^2 1
Genus Crypturgus *87A*
^4 1
funicle 5-segmented; sutures of antennal club not fully distinguishable:
^5 1
Genus Pseudothamnurgus *95A*
#27D F56-57.A12 backto #25B
^1 1
pronotum with uniformly distributed granules (fig. 56) *29A*
^3 1
pronotum punctate or smooth, anterior 1/2 granulate (fig. 57) *29B*
#29A F58-59.A12 backto #27D
^4 35
eyes anteriorly slightly emarginate (fig. 58);
^5 35
pronotum rough sculptured;
^5 35
body with dense, long, filiform setae:
^6 35
Genus Dryocoetes *91A*
^19 35
eyes bipartite (fig. 59); pronotum with
^20 35
fine, oblique granules; elytral setae
^21 35
sparse: Genus Trypodendron *161A*
#29B F60-61.A12 backto #27D
^1 1
posterior ventral elytral margin directly surrounding abdomen (fig. 60) *29C*
^3 1
posterior ventral elytra with 2 margins: inner surrounding abdomen, outer
^4 1
defining elytral declivity (fig. 61) *31C*
#29C backto #29B
^1 1
anterior pronotal margin smooth *29D*
^3 1
anterior pronotal margin with fine granules *29E*
#29D F62-63.A12 backto #29C
^2 2
scutellum small, indistinct (fig. 62): Genus Xylocleptes *93A*
^18 2
scutellum visible (fig. 63): Genus Lymantor *97A*
#29E backto #29C
^1 1
posterior pronotal margin bordered *31A*
^3 1
posterior pronotal margin not bordered *31B*
#31A F64-65.A12 backto #29E
^1 1
antennal club with 2 bowed sutures (fig. 64); elytral declivity with shallow
^2 1
furrow adjacent suture: Genus Gnathotrichus *123A*
^4 1
antennal club laterally, deeply indented (fig. 65); elytral declivity with
^5 1
deep, broad furrow adjacent suture:
^6 1
Genus Pityophthorus *115A*
#31B backto #29E
^1 1
medial posterior pronotum with flat, longitudinal ridge; elytral declivity
^2 1
with spines, smaller in female: Genus Pityogenes *129A*
^4 1
pronotum without flat, longitudinal ridge; elytral declivity without spines,
^5 1
only with granules: Genus Taphrorychus *125A*
#31C F66.A12 backto #29B
^1 1
2nd spine on elytral declivity cone-shaped *31D*
^3 1
2nd spine on elytral declivity distally hooked (fig. 66):
^4 1
Genus Pityokteines *137A*
#31D F67-68.A12 backto #31C
^1 43
elytral declivity regularly rounded
^2 43
(fig. 67); 1st interstria on declivity
^3 43
with row of granules; sutures of club
^4 48
bowed: Genus Ips *147A*
^17 44
elytral declivity sloping steeply
^18 44
downward (fig. 68); 1st interstria on
^19 44
declivity without rows of granules;
^20 49
sutures of club straight:
^21 49
Genus Orthotomicus *141A*
#105A backto #25E
^1 1
Cryphalus: 1.1 to 2 mm; eyes slightly anteriorly emarginate; funicle 4-seg-
^2 1
mented; pronotum arched; anterior medial pronotal margin without protruding
^3 1
granules; posterior pronotal margin with fine, ridged border; elytral declivity
^4 1
rounded; monogamous; host: confers.
^6 1
elytra with sparse, long, filiform setae *105B*
^8 1
elytra with rows of short, filiform or squamiform setae *105C*
#105B F177.A12 backto #105A
^1 1
pronotum anteriorly constricted (fig. 177); lateral pronotum with short,
^2 1
filiform setae; elytra 1.75 x as long as wide; with rows of filiform setae
^3 1
and indistinct striae; body dark brown; antennae and legs light brown;
^4 1
1.1 to 1.8 mm: Cryphalus piceae RATZEBURG *
^6 1
pronotum round, arched, medial anterior pronotum with tubercles; lateral
^7 1
pronotum and head with sparse, long, filiform setae; elytra only 1.5 x as long
^8 1
as wide, with rows of dense, long, filiform setae; body dark brown; antennae
^9 1
and legs light brown; 1.2 to 1.8 mm: Cryphalus numidicus EICHHOFF *
#105C F178.A12 backto #105A
^21 1
elytra 1.33 to 1.36 x as long as wide; body cylindrical and broadened (fig.
^22 1
178); frons with fine granules and short, filiform setae; pronotum with dense,
^23 1
fine punctures; elytra with fine striae; body dark brown, dull, with dense,
^24 1
short, filiform setae; antennae and legs light brown; 2 mm:
^25 1
Cryphalus intermedius FERRARI *
^27 1
elytra 1.5 to 1.7 x as long as wide; body cylindrical and elongated *107A*
#107A F178A.A12 F179.A12 backto #105C
^16 1
tubercles on pronotum large (fig. 178A); elytra 1.6 to 1.67 x as long as wide,
^17 1
with fine, indistinct striae; interstriae with oval, light brown, filiform
^18 1
setae; body black, dull; elytra black to brown; antennae and legs light brown;
^19 1
1.5 to 2 mm: Cryphalus saltuarius WEISE *
^21 1
tubercles on pronotum find (fig. 179); elytra with distinct striae; elytra
^22 1
1.5 to 1.57 x as long as wide, with distinct striae; frons with thin, short,
^23 1
filiform setae; pronotum with short, filiform setae; interstriae with fine,
^24 1
squamiform setae; body dark brown to black, dull, antennae and legs light
^25 1
brown; 1.2 to 1.7 mm: Cryphalus abietis RATZEBURG *
#113A F183B.A12 F183A.A12 backto #25E
^1 1
Hypothenemus: anterior pronotum with tubercles; club long, oval, with 3
^2 1
slightly, laterally indented sutures; female funicle 4-segmented (fig. 183A);
^3 1
male funicle 3-4 segmented; male often smaller than female; monogamous;
^4 1
polyphagous.
^6 1
anterior pronotal margin granulate (fig. 183B); elytra 2 x as long as wide,
^7 1
with distinct striae; interstriae with rows of short, thick, squamiform setae;
^8 1
body brown to black; antennae and legs light brown; 0.7 to 1.3 mm:
^9 1
Hypothenemus eruditus WESTWOOD *
#109A F180.A12 F181-182.A12 backto #25F
^1 1
Ernoporus: 1.5 to 2 mm; funicle 4-segmented; club oval, with 3 bowed sutures
^2 1
(fig. 180); pronotum arched; medial antennal pronotal margin with 2 protruding
^3 1
tubercles; anterior pronotum with irregularly distributed tubercles;
^4 1
monogamous; host: deciduous trees.
^6 1
elytra 2 x as long as wide, with fine striae and rows of fine, light brown,
^7 34
squamiform setae and short, thick, filiform
^8 41
setae (fig. 181); anterior pronotum
^9 41
constricted, medial anterior pronotum
^10 41
with irregularly distributed tubercles;
^11 41
frons with filiform setae and granules;
^12 41
body black; pronotum and lateral elytra
^13 41
reddish; 1.5 to 2 mm:
^14 41
Ernoporus caucasicus LINDEMANN *
^17 41
elytra 2.5 x as long as wide with fine
^18 41
white to light brown squamiform setae
^19 41
(fig. 182); striae indistinct; anterior
^20 41
pronotum only slightly constricted
^21 41
with irregularly distributed tubercles;
^22 41
granulate frons with medial depression
^23 41
and fine, filiform setae; body black,
^24 41
shiny; antennae and legs brown to light
^25 41
brown; 1.5 to 1.8 mm:
^26 41
Ernoporus fagi FABRICIUS *
#111A F183.A12 backto #25F
^1 1
Cryphalops: funicle 4-segmented; club oval with 3 straight sutures; pronotum
^2 1
arched; medial anterior pronotal margin with 4 tubercles; medial anterior
^3 1
1/2 pronotum with 4 rows of tubercles, posteriorly decreasing in width;
^4 1
monogamous; host: deciduous trees.
^6 1
elytra at most 2 x as long as wide with small, thick, filiform setae (fig. 183);
^7 1
body brown; 1.1 to 1.5 mm: Cryphalops tiliae PANZER *
#99A F172A-B.A12 backto #27A
^1 1
Trypophloeus: 1.3 to 2.2 mm; funicle 5-segmented; club long, conical, 3-sutured;
^2 1
elytra with squamiform setae; sexual dimorphism: male elytral declivity long,
^3 1
flat, sometimes pointed tubercle on 4th interstria; female elytral declivity
^4 1
short, flat; monogamous; host: deciduous trees.
^6 1
elytra at most 2 x as long as wide *99B*
^8 1
elytra 2.5 x as long as wide, with sparse, fine, filiform setae (fig. 172A);
^9 1
fine striae reach posterior elytral margin; interstriae with rows of short,
^10 1
white filiform setae; declivity slightly flattened (fig. 172B), suture slightly
^11 1
indented; granules on declivity absent; pronotum not longer than wide, finely
^12 1
wrinkled, dull; frons with medial, longitudinal furrow; body dark brown,
^13 1
antennae and legs light brown; 1.5 mm: Trypophloeus alni LINDEMANN *
#99B F172C.A12 backto #99A
^1 1
elytra 1.5 to 1.57 x as long as wide; male tubercle on 4th interstria on
^2 1
declivity larger than in female *99C*
^4 1
elytra 1.62 to 1.65 x as long as wide, lateral elytra with striae (fig. 172C);
^5 1
male granule on 4th interstria on declivity hardly visible in female; body
^6 1
black; antennae and legs light brown; club and femur brown; 1.5 to 1.8 mm:
^7 1
Trypophloeus rybinskii REITTER *
#99C backto #99B
^1 1
posterior pronotum fine sparsely punctate; lateral pronotum regularly
^2 1
punctured *101A*
^4 1
posterior pronotum densely punctured; elytral declivity rounded; elytra with
^5 1
striae *101C*
#101A F173-174.A12 backto #99C
^1 1
interstriae on elytral declivity without squamiform setae; suture on elytral
^2 1
declivity not indented *101B*
^4 1
interstriae on declivity with squamiform setae; suture on elytral declivity
^5 1
slightly indented (fig. 173); elytra without striae, nearly smooth, only with
^6 1
rows of filiform setae; setae not on granules (fig. 174); male declivity flat,
^7 1
covering posterior 1/2 of elytra;
^8 1
body dark brown; 1.2 to 1.6 mm:
^9 1
Trypophloeus asperatus GYLLENHAL *
#101B backto #101A
^1 1
frons flat, wrinkled; pronotum wider than long; anterior pronotal margin with
^2 1
4 long and 2 small tubercles; male elytral declivity with lateral tubercle;
^3 1
body reddish brown, shiny; antennae and tarsi light brown; 1.4 to 1.5 mm:
^4 1
Trypophloeus berezinae STARK *
^6 1
frons round, fine densely punctate; pronotum as wide as long; anterior
^7 1
pronotal margin with 6 tubercles; male elytral declivity on 4th interstria
^8 1
with small tubercle, absent in female; body black, shiny; 1.5 to 2.1 mm:
^9 1
Trypophloeus spiculatus EGGERS *
#101C backto #99C
^1 1
posterior pronotum densely punctured, rough, dull; elytra 1.5 x as long as
^2 1
wide, with striae; male tubercle of 4th interstria on declivity smaller in
^3 1
female; declivity rounded; body black; antennae and legs light brown; 1.7 to
^4 1
2.1 mm: Trypophloeus tremulae STARK *
^6 1
pronotum shiny, roughly punctured posteriorly; elytra 1.55 to 1.57 x as long
^7 1
as wide *103A*
#103A F175-176.A12 backto #101C
^1 1
elytra with striae and rows of filiform setae (fig. 175); male pointed tubercle
^2 1
of 4th interstria on declivity smaller in female; declivity steep, covering
^3 1
posterior 1/3 elytra, with setae on granules (fig. 176); body dark brown to
^4 1
black; antennae and legs light brown; 1.5 to 2.1 mm:
^5 1
Trypophloeus granulatus RATZEBURG *
^7 1
elytra with striae and long squamiform setae; elytral suture ridged on
^8 1
declivity; medial anterior pronotal
^9 1
margin with 4 tubercles; male with
^10 1
3 thin, pointed tubercles on 4th
^11 1
interstria on declivity, absent in
^12 1
female; body black; 1.4 to 2 mm:
^13 1
Trypophloeus bispinulus EGGERS *
#159A F255.A12 backto #27B
^1 1
Xylosandrus: 1st and 5th funicle segments with similar width; prothoracic
^2 1
coxae separated by broad prosternal process; lateral declivital margin sharp;
^3 1
female pronotum rounded, arched, smooth, anterior 1/2 with granules; male
^4 1
lighter colored than female; wings rudimentary or absent; incapable of flying;
^5 1
polyphagous
^6 1
female 2 to 2.5 mm (fig. 255); elytra with regular striae; declivity oval,
^7 1
flat, lateral margin from 7th interstria; body dark brown to black, shiny;
^8 1
male 1 to 1.8 mm; body long, light brown, seldom darker; lateral declivity
^9 1
with fine margin; granules on interstriae of declivity with small, filiform
^10 1
setae: Xylosandrus germanus BLANDFORD *
^26 1
female 1.5 to 1.8 mm; body short, cylindrical, reddish yellow, shiny; anterior
^27 1
1/2 pronotum granulate with filiform setae, posterior 1/2 pronotum and scutellum
^28 1
smooth; fine striae on elytra more distinct on rounded declivity; elytra
^29 1
anteriorly raised: Xylosandrus morigerus BLANDFORD *
#153A F245.A12 backto #27B
^1 1
Xyleborus: female 2.1 to 3.8 mm; body reddish brown to black; male 1.5 to 3 mm;
^2 1
light brown to reddish brown; prothoracic coxae separated by broad prosternal
^3 1
process; pronotum flat; elytra flat; wings rudimentary,incapable of flying;
^4 1
polyphagous.
^22 1
female pronotum slightly arched, longer than wide, posteriorly smooth (fig.
^23 1
245); elytral declivity dull, with fine row of granules adjacent suture and
^24 1
on 3rd interstria; body dark brown; head and pronotum brown; antennae and legs
^25 1
light brown to yellow; 2 to 2.4 mm; Male pronotum longer than wide, slightly
^26 1
rounded, anterior 1/2 with tubercles; pronotum and elytra with fine filiform
^27 1
setae; elytral declivity punctured; 1.5 - 1.8 mm: Xyleborus saxeseni RATZEBURG *
^29 1
female pronotum arched *153B*
#153B backto #153A
^1 1
female pronotum longer than wide, cylindrical, and anteriorly raised; male's
^2 1
pronotum anteriorly depressed, anterior pronotal margin with raised border
^3 1
sometimes medially protruding *153C*
^5 1
female pronotum not longer than wide *157A*
#153C F246-247.A12 backto #153B
^1 1
female pronotum nearly square, anterior pronotal margin medially slightly
^2 1
emarginate (fig. 246); elytra with striae; interstriae with sparse punctures
^3 1
and granules on declivity; body dark brown; elytra brown; 2.5 to 4 mm; male
^4 1
frons with medial longitudinal ridge; anterior pronotum depressed, round, with
^5 1
rough, dense punctures; 3 mm: Xyleborus eurygraphus RATZEBURG *
^7 1
female anterior pronotum constricted (fig. 247) *155A*
#155A F248-249.A12 backto #153C
^1 1
female anterior pronotum rounded, medially raised (fig. 248); suture depressed
^2 1
on declivity; granules on 1st, 3rd, 4th and 5th interstriae on declivity;
^3 1
body brown; punctures, suture, and scutellum red; antennae and legs light
^4 1
brown to yellow; 2.1 to 3.5 mm; male frons without ridge; anterior 1/2 pronotum
^5 1
deeply depressed, margin medially protruding and pointed; body light brown;
^6 1
antennae and legs yellow; 2.1 to 2.7 mm: Xyleborus pfeili RATZEBURG *
^8 1
female pronotum anteriorly raised *155B*
#155B F250-251.A12 F248-249.A12 backto #155A
female anterior pronotal margin rounded (fig. 249); body cylindrical; elytral
^2 1
suture not ridged; 3rd interstria with 6 conical tubercles; declivity flat,
^3 1
dull, without striae (fig. 250); body light brown; 3 to 3.5 mm; male anterior
^4 1
1/2 pronotum deeply depressed, granulate; anterior pronotal margin medially
^5 1
protruding into long pointed tubercle; body light brown; 2 to 2.5 mm:
^6 1
Xyleborus monographus FRABRICIUS *
^8 1
female elytra with regular, dark striae, posterior slightly constricted;
^9 1
declivity steep, shiny (fig. 251); suture depressed; interstriae with rows of
^10 1
granules on declivity; body reddish brown; 2 to 2.7 mm; male anterior 1/2
^11 1
pronotum depressed, anterior pronotal margin medially protruding into long
^12 1
pointed tubercle; body reddish brown; 1.9 to 2 mm:
^13 1
Xyleborus dryographus RATZEBURG *
#157A F252-253.A12 backto #153B
^1 1
female posterior pronotum raised (fig. 252); pronotum punctate, anterior 1/2
^2 1
with granules (fig. 253); elytra and interstriae with fine rows of punctures;
^3 1
declivity rounded, adjacent suture with shallow furrow; small filiform setae
^4 1
on granules of interstriae of declivity; body brown; antennae and legs light
^5 1
brown; 2.3 to 3.4 mm; Male pronotum longer than wide, smooth; elytra as flat
^6 1
as pronotum, with sparse, filiform setae; metathoracic tibiae distally produced
^7 1
into long, straight spine; body light brown; 1.5 mm (male, see above):
^8 1
Xyleborus cryptographus RATZEBURG *
^9 1
female pronotum medially raised; anterior 1/2 pronotum with concentric granules;
^10 1
posterior pronotum smooth (fig. 254); elytra with rough striae; declivity with
^11 1
rows of granules and furrow adjacent suture; body dark brown, shiny, with
^12 1
sparse, filiform setae; antennae and legs light brown; 3 to 3.8 mm; male
^13 1
pronotum longer than wide, heart-shaped, flat; anterior 1/2 pronotum with fine
^14 1
granules; posterior elytra wider, slightly rounded, with long, filiform setae;
^15 1
declivity covering posteriot 1/2 of elytra; body brown; 1.8 to 2.4 mm (male):
^16 31
Xyleborus dispar FABRICIUS *
#95A F170A.A12 backto #27C
^1 1
Pseudothamnurgus: 1.2 to 2 mm; male frons with medial, transverse indentation;
^2 1
funicle 5-segmented; club flat, oval, 2-sutured; pronotum longer than wide,
^3 1
punctate, with medial longitudinal band without punctures; declivity flat,
^4 1
punctate; suture ridged on declivity; 1st stria adjacent suture extends
^5 1
posterior only 1/2 length of elytra; antennae and legs light brown.
^7 1
pronotum roughly punctured, with medial, longitudinal ridged band without
^8 1
punctures; punctures of striae dense; female frons granulate; body dark brown;
^9 1
elytra black; 1.2 to 2 mm: Pseudothamnurgus mediterraneus EGGERS *
^11 1
pronotum fine, sparsely punctured with medial, longitudinal band without
^12 1
punctures (fig. 170A); punctures of striae fine, irregularly distributed;
^13 1
elytra posteriorly wider; body reddish brown; 1.2 to 1.8 mm:
^14 1
Pseudothamnurgus scrutator PANDELLE *
#91A F164.A12 backto #29A
^1 1
Dryocoetes: 2 to 4.5 mm; funicle 5-segmented (fig. 164); eyes laterally
^2 1
emarginate; pronotum not arched, anterior 1/2 pronotum roughly sculptured,
^3 1
posterior 1/2 pronotum finely punctured; elytra with striae; declivity rounded;
^4 1
monogamous; polyphagous.
^6 1
furrow on elytral declivity absent *91B*
^8 1
elytral declivity with furrow adjacent suture *91C*
#91B F165-166.A12 backto #91A
^13 1
pronotum anteriorly and posteriorly constricted (fig. 165); punctures of striae
^14 1
large on declivity; interstriae with fine punctures and rows of long, light
^15 1
brown, filiform setae; body dark brown to reddish brown; 3 to 4.3 mm:
^16 1
Dryocoetes autographus RATZEBURG *
^18 1
pronotum anteriorly constricted (fig. 166); body reddish brown; 3 to 3.2 mm:
^19 1
Dryocoetes baicalicus REITTER *
#91C F167-168.A12 backto #91A
^12 1
pronotum slightly longer than wide (fig. 167); elytral suture on declivity with
^13 1
row of granules; male frons almost bare; female frons with dense, light brown,
^14 1
filiform setae; body reddish brown; antennae and legs light brown; 2.5 to 3.3
^15 1
mm: Dryocoetes villosus FABRICIUS *
^17 1
pronotum as long as wide (fig. 168); granules on elytral suture absent; elytra
^18 1
with dense, rough striae; interstriae with fine row of punctures; body brown
^19 1
with sparse, filiform setae; elytra reddish brown; antennae and legs light
^20 1
brown; 2 to 2.6 mm: Dryocoetes alni GEORG *
#161A F256-257.A12 F258-261.A12 bacto #29A
^1 1
Trypodendron: 2.8 to 3.8 mm; body cylindrical; eyes bipartite; funicle
^2 1
4-segmented; club without distinct sutures; female pronotum rounded, anterior
^3 1
1/2 with granules; frons rounded (fig. 256); male pronotum flat, anterior 1/2
^4 1
with granules; frons concave (fig. 257); elytra with striae; declivity
^5 1
covering less than posterior 1/2 of elytra, shallow furrow adjacent suture;
^6 1
filiform setae of pronotum longer than elytral setae; monogamous; ambrosia
^7 1
beetles; polyphagous.
^9 1
short, broad furrow adjacent declivital suture (fig. 258); club distal and
^10 1
laterally pointed (fig. 259); pronotum black; elytra light brown to yellow
^11 1
with medial and lateral black bands (fig. 260); 3 to 3.5 mm:
^12 1
Trypodendron domesticum L. *
^14 1
elytral declivity with short, filiform setae and shallow furrow adjacent suture
^15 1
(fig. 261) *161B*
#161B F262-264.A12 F260-265.A12 backto #161A
^1 1
club oval (fig. 262); striae of elytra finer on declivity; declivity dull, with
^2 1
sparse, filiform setae (fig. 263); pronotum reddish brown; elytra light brown
^3 1
with longitudinal, black bands; 2.8 to 3.8 mm: Trypodendron lineatum OLIVER *
^5 1
distal club laterally rounded (fig. 264); elytra with rough striae; declivity
^6 1
wrinkled with shallow furrow adjacent suture; body light brown, shiny, with
^7 1
longitudinal black bands and short filiform setae (fig. 265); 3.2 to 3.8 mm:
^8 1
Trypodendron signatum FABRICIUS *
#93A F170.A12 F169.A12 backto #29D
^1 1
Xylocleptes: 1.5 to 3.4 mm; funicle 5-segmented; club flat, round, with bowed
^2 1
sutures (fig. 169); anterior pronotum granulate, posterior pronotum roughly
^3 1
punctured; elytral declivity with shiny furrow; male margins of furrow with
^4 1
tubercles, female with granules; polygamous; host: Clematis.
^6 1
male elytral declivity round, with lateral, pointed tubercle; female declivity
^7 1
with furrow and row of granules; elytral suture ridged and granulate (fig.
^8 1
170); male posterior elytral margin emarginate; body brown, shiny, with grey,
^9 1
filiform setae; legs reddish brown; antennae and tarsi light brown; 2.3 to 3.4
^10 1
mm: Xylocleptes bispinus DUFTSCHMIDT *
^26 1
male elytral declivity with lateral, pointed, conical tubercle; female elytral
^27 1
suture smooth, declivity with narrow furrow adjacent suture; posterior elytral
^28 1
margin not emarginate; body black; antennae and legs reddish brown; 1.5 to 2 mm:
^29 1
Xylocleptes biuncus REITTER *
#97A F172.A12 F171.A12 backto #29D
^1 1
Lymantor: 1.6 to 2.2 mm; funicle 4-segmented (fig. 171); elytra with dense
^2 1
striae; declivity with smooth furrow adjacent suture; male frons with large,
^3 1
shiny tubercle; monogamous; host deciduous trees.
^5 1
declivity flat, with slight, smooth, and shiny furrow adjacent suture (fig.
^6 1
172); body dark brown; legs light brown; 1.7 to 2.2 mm: Lymantor coryli PERRIS *
^8 1
declivity rounded, with striae, and furrow adjacent suture; 1.6 to 2 mm:
^9 1
Lymantor aceris LINDEMANN *
#123A F195.A12 F194.A12 backto #31A
^1 1
Gnathotrichus: funicle 5-segmented; club flat, with 2 bowed sutures (fig. 194);
^2 1
male frons without setae; female frons wrinkled, with filiform setae; pronotum
^3 1
large, longer than wide; posterior pronotum wider than anterior elytra;
^4 1
anterior pronotum with granules; posterior pronotum with fine punctures;
^5 1
medial pronotum with depression; body cylindrical; elytral declivity rounded,
^6 1
with longitudinal furrow adjacent suture; striae and interstriae with sparse,
^7 1
fine punctures and long, thin filiform setae; monogamous; host: conifers.
^9 1
body long, cylindrical; pronotum 0.75 x as long as elytra (fig. 195); elytra
^10 1
with fine striae; granules on margin of furrow with 3 long filiform setae;
^11 1
body dark brown; elytra reddish brown; antennae light brown; 3.2 to 3.5 mm:
^12 1
Gnathotrichus materiarius FITCH *
#115A F184.A12 backto #31A
^1 1
Pityophthorus: 0.9 to 2.5 mm; body cylindrical; funicle 5-segmented, 1st
^2 1
segment alone as long as all the others; club 1.5 x longer than wide, with
^3 1
3 laterally indented sutures; eyes anteriorly slightly emarginate; pronotum
^4 1
longer than wide, anteriorly with concentric tubercles; posterior pronotum
^5 1
punctate, with fine border; declivity adjacent suture with furrow; polygamous;
^6 1
host: conifers.
^8 1
margins of furrow on declivity with long, filiform setae on granules *115B*
^10 1
margins of furrow on declivity without long, filiform setae (fig. 184) *119A*
#115B F185-186.A12 backto #115A
^8 1
posterior elytral margin protruding
^9 1
beyond abdomen (fig. 185); elytra
^10 1
2.5 x as long as wide; body long,
^11 1
cylindrical *115C*
^21 1
posterior elytral margin rounded,
^22 1
not protruding (fig. 186) *117A*
#115C backto #115B
^1 1
posterior 1/3 of elytra with furrow *115D*
^3 1
posterior 1/2 of elytra with furrow; elytra with fine striae; granulated
^4 1
elytral suture depressed within lateral margins of furrow; body reddish brown;
^5 1
1.1 to 1.7 mm: Pityophthorus exculptus RATZEBURG *
#115D F187-188.A12 backto #115C
^1 1
female frons with dense, long, light brown, filiform setae (fig. 187); elytra
^2 1
with rough striae (fig. 188); suture and margins of furrow on declivity with
^3 1
fine granules and filiform setae; body dark brown; antennae and tarsi brown;
^4 1
1.1 to 1.5 mm: Pityophthorus pityographus RATZEBURG *
^11 1
female frons with crown of light brown,
^12 1
filiform setae; elytral suture and
^13 1
margins of furrow with fine granules
^14 1
and filiform setae; body dark brown;
^15 1
antennae and legs brown;
^16 1
1.3 to 2.5 mm:
^17 1
Pityophthorus micrographus L. *
#117A backto #115B
^1 1
elytral declivity with broad, smooth furrow *117B*
^3 1
elytral declivity with narrow, reticulate furrow *117C*
#117B F189-190.A12 backto #117A
^13 1
Elytral suture without granules and not depressed within lateral margins of
^14 1
furrow (fig. 189); furrow with row of filiform setae on small granules; male
^15 1
frons punctate; female frons with crown of blond filiform setae; body black
^16 1
to brown; 1.6 to 2.2 mm: Pityophthorus lichtensteini RATZEBURG *
^17 5
Elytral suture depressed within lateral margins of furrow (fig. 190); body
^18 1
dark brown; 2.5 to 2.8 mm: Pityophthorus knoteki REITTER *
#117C backto #117A
^1 1
elytra with dense, rough, irregular striae; furrow on declivity reticular,
^2 1
dull; pronotum with lateral tubercle; body black to brown, shiny; 1 to 1.2 mm:
^3 1
Pityophthorus pubescens MARSHAM *
^5 1
elytra with fine regular striae; furrow on declivity smooth, shiny; medial
^6 1
anterior pronotal margin with 3 to 4 tubercles; posterior pronotum rough,
^7 1
densely punctured, with medial, longitudinal shiny band without punctures;
^8 1
male frons dense, roughly punctured, with filiform setae and medial,
^9 1
longitudinal ridge; female frons with long, blond, filiform setae; body dark
^10 1
brown; antennae and legs light brown; 1.1 to 1.5 mm:
^11 1
Pityophthorus cephalonicae PFEFFER *
#119A backto #115A
^1 1
elytral declivity with sparse, short, filiform setae; furrow on declivity
^2 1
broad and smooth *121A*
^4 1
elytral declivity with dense, short, filiform setae; furrow on declivity
^5 1
narrow *119B*
#119B F191.A12 backto #119A
^17 1
elytra 1.5 x as long as wide, with irregular striae (fig. 191); frons roughly
^18 1
punctured; female frons with short, medial, longitudinal ridge; body dark
^19 1
brown; antennae and legs reddish brown; 1.1 to 1.5 mm:
^20 1
Pityophthorus carniolicus WICHMANN *
^23 1
elytra 1.75 to 1.85 x longer than wide, with irregular striae *119C*
#119C F192.A12 backto #119B
^1 1
frons roughly punctured, with short, medial, longitudinal ridge; pronotum with
^2 1
narrow, medial, longitudinal band without punctures; body dark brown, smooth;
^3 1
antennae and legs light brown; 1.3 to 1.5 mm:
^4 1
Pityophthorus morosovi SPESSIVTSEFF *
^6 1
frons with medial, longitudinal ridge; pronotum with broad, medial,
^7 1
longitudinal band without punctures (fig. 192); body dark brown, smooth;
^8 1
antennae and legs light brown; 1.7 to 1.9 mm: Pityophthorus henscheli SEITNER *
#121A backto #119A
^1 1
elytra 1.7 x as long as wide, with fine, regular striae; interstriae slightly
^2 1
reticular *121B*
^4 1
elytra 1.7 to 1.8 x as long as wide, anterior 1/2 and lateral areas irregularly
^5 1
punctured; interstriae reticular, dull; pronotum fine, densely punctate *121C*
#121B F193.A12 backto #121A
^17 1
male frons without setae, punctate, rounded, with narrow, medial, longitudinal
^18 1
band without punctures; female frons with medial semicircular indentation and
^19 1
lateral brushes of long blond filiform setae above bases of mandibles; pronotum
^20 1
densely punctured, shiny (fig. 193); body black; antennae and legs light brown;
^21 1
1.5 to 2.2 mm: Pityophthorus glabratus EICHHOFF *
^24 1
frons round, roughly punctate, with short, medial, longitudinal ridge and
^25 1
short filiform setae; pronotum dull; body dark brown; antennae, pronotum and
^26 1
legs light brown; 1.5 to 1.7 mm: Pityophthorus tr„g†rdhi SPESSIVTSEFF *
#121C backto #121A
^1 1
elytral declivity with shiny, laterally granulated furrow; male frons round,
^2 1
with medial, longitudinal ridge; female frons with crown of dense, long, blond,
^3 1
filiform setae; 1.9 mm: Pityophthorus balcanicus PFEFFER *
^5 1
elytral declivity with dull, reticular, laterally granulated furrow; elytra
^6 1
with fine, irregular striae; male frons rounded, with distinct, medial,
^7 1
longitudinal ridge; female frons flat, with sparse, filiform setae; body black;
^8 1
elytra dark brown; antennae and legs reddish brown; tarsi reddish yellow;
^9 1
1.5 to 2.3 mm: Pityophthorus buyssoni REITTER *
#129A backto #31B
^1 1
Pityogenes: 1.5 to 3.5 mm; body stocky; funicle 5-segmented; pronotum large
^2 1
relative to elytra, posterior pronotum punctate, with medial, longitudinal
^3 1
ridged band without punctures; elytral declivity rounded; male lateral
^4 1
declivity with spines, female with tubercles; polygamous; host: conifers.
^6 1
male elytral declivity with 3 conical spines; female frons with round, medial
^7 1
depression; male frons with medial tubercle *129B*
^9 1
male elytral declivity with tubercles and 1 to 2 dorsal spines; female frons
^10 1
without depression *131B*
#129B F200.A12 backto #129A
^1 1
elytral striae laterally indistinct; rounded elytral declivity covering post-
^2 1
erior 1/2 elytra; male declivity with 3 conical spines (fig. 200); female with
^3 1
3 tubercles; body black; elytra reddish brown, shiny; 1.6 to 2.9 mm:
^4 1
Pityogenes chalcographus L. *
^6 1
elytral striae distinct to posterior elytral margin; distance between 1st and
^7 1
2nd conical spines less than distance between 2nd and 3rd spines *129C*
#129C F201-202.A12 backto #129B
^1 1
punctures of elytral interstriae finer
^2 1
than those of striae *131A*
^5 1
punctures of elytral interstriae and striae
^6 1
similar (fig. 201); elytra roughly
^7 1
punctured; pronotum nearly square (fig. 202);
^8 1
2 to 2.4 mm:
^9 1
Pityogenes saalasi EGGERS *
#131A F203.A12 F204.A12 backto #129C
^18 1
distance between 2nd and 3rd conical spines on elytral declivity 1.5 x longer
^19 1
than distance between 1st and 2nd conical spines; elytra with filiform setae;
^20 1
pronotum anteriorly constricted (fig. 203); body black; elytra reddish brown,
^21 1
slightly shiny; 2.2 to 2.5 mm: Pityogenes trepanatus N™RDLINGER *
^24 1
distance between 1st and 2nd conical spines on elytral declivity 0.5 x as long
^25 1
as distance between 2nd and 3rd conical spines (fig. 204); elytra 1.45 to 1.55
^26 1
x as long as wide; posterior elytra with fine striae, shiny; 1.8 to 2.3 mm:
^27 1
Pityogenes irkutensis EGGERS *
#131B F205.A12 F206.A12 backto #129A
^21 1
dorsal declivity adjacent suture with small granule; lateral margin of
^22 1
declivity with 1 hooked spine and ventral granules (fig. 205); declivity shiny
^23 1
(fig. 206); female frons with small, bilateral indentations and medial triangle
^24 1
of short, filiform setae; body black to brown, shiny; 2 to 2.8 mm:
^25 1
Pityogenes bidentatus HERBST *
^27 1
lateral margin of elytral declivity with 1 to 2 distinct spines and ventral
^28 1
granules *131C*
#131C backto #131B
^1 1
1st spine adjacent suture on elytral declivity distinct *133A*
^3 1
1st spine adjacent suture on elytral declivity indistinct; 2nd spine thick,
^4 1
hooked, and medially oriented; outer declivital margin with granule between
^5 1
2nd spine and conical tubercle; declivity with long, filiform setae; 2.5 mm:
^6 1
Pityogenes pennidens REITTER *
#133A backto #131C
^1 1
anterior elytra up to the 1st spine adjacent suture on declivity as long as
^2 1
wide *133B*
^4 1
anterior elytra up to the 1st spine adjacent suture on declivity wider than
^5 1
long *135A*
#133B F207.A12 backto #133A
^23 1
1st spine adjacent suture on elytral declivity small, distance to 2nd spine
^24 1
short; conical tubercle on declivital margin thick and 1/2 as long as spine
^25 1
(fig. 207); male declivity rounded; female frons with long, blond, filiform
^26 1
setae; body black to brown; elytra reddish brown; antennae and legs light
^27 1
brown; 1.6 to 2.4 mm: Pityogenes quadridens HARTIG *
^29 1
1st spine adjacent suture on male elytral declivity 1/2 as long as thick,
^30 1
pointed 2nd spine; distance to 2nd spine long *133C*
#133C backto #133B
^1 1
2nd spine on elytral declivity hooked and pointed; filiform setae on granule
^2 1
between 2nd spine and conical tubercle; lateral declivital margin with sparse,
^3 1
long, filiform setae; female elytra reddish brown; male elytra dark brown;
^4 1
body black to brown; antennae and legs light brown; 2.2 to 2.8 mm:
^5 1
Pityogenes conjunctus REITTER *
^7 1
2nd spine on elytral declivity thick, bifurcate; male declivity rounded, dull;
^8 1
frons rounded, with sparse, filiform setae; body brown; antennae and legs
^9 1
light brown; 2 to 3 mm: Pityogenes herbellae STROHMEYER *
#135A F208-210.A12 backto #133A
^1 1
indistinct granule between 2nd spine and
^2 1
conical tubercle of elytra; declivity
^3 1
without filiform setae (fig. 208); female
^4 1
frons with long, blond, filiform setae;
^5 1
female declivity reddish brown; body black
^6 1
to brown; antennae light brown; legs
^7 1
reddish brown; 1.8 to 2.6 mm:
^8 1
Pityogenes bistridentatus EICHHOFF *
^10 1
2 small tubercles betwen 2nd spine
^11 1
and conical tubercle of elytra (figs.
^12 1
209, 210); female frons with medial
^13 1
triangle of short, filiform setae
^14 1
and adjacent shallow furrows; body
^15 1
light brown; 1.8 to 2.5 mm:
^16 1
Pityogenes calcaratus EICHHOFF *
#125A F196.A12 backto #31B
^1 1
Taphrorychus: 1.5 to 2.6 mm; funicle 5-segmented; antennal club flat, round,
^2 1
with 3 slightly bowed sutures (fig. 196); pronotum as long as wide, anteriorly
^3 1
constricted, with rows of fine granules; elytra with striae; male elytral
^4 1
declivity flat; female declivity rounded; male frons granulate and with sparse
^5 1
filiform setae; female frons with crown of dense, short, blond, filiform setae;
^6 1
body black to brown; antennae and legs light brown; polygamous; deciduous trees.
^8 1
elytral striae distinct only laterally *125B*
^10 1
elytra shiny, striae dense *127A*
#125B F197.A12 backto #125A
^17 1
pronotum longer than wide, anterior with granules, posterior with dense, fine
^18 1
punctures (fig. 197); reticular elytra as wide as pronotum, 1.9 x as long as
^19 1
wide; declivity steep with fine, squamiform, and filiform setae and with 3 rows
^20 1
of granules; striae absent; body dull; 1.6 to 2 mm:
^21 1
Taphrorychus hirtellus EICHHOFF *
^24 1
pronotum anterior 1/2 with granules, posterior 1/2 roughly punctured; body
^25 1
shiny, with fine irregular punctures and filiform setae; elytra 1.7 x longer
^26 1
than wide; 1.8 to 2.4 mm: Taphrorychus minor EGGERS *
#127A F198.A12 backto #125A
^18 1
elytral declivity adjacent suture with 5 or 6 tubercles; 3rd and 5th
^19 1
interstriae with 3 to 5 fine tubercles; rounded declivity indistinctly
^20 1
punctured (fig. 198); female declivity dull, with slight furrow adjacent suture
^21 1
and with 3 rows of indistinct granules; body black to brown, shiny; antennae
^22 1
and legs light brown; 1.8 to 2.3 mm: Taphrorychus villifrons DUFOUR *
^25 1
suture on elytral declivity with 8 to 11 tubercles *127B*
#127B backto to #127A
^1 1
suture and interstriae on elytral declivity with 8 or 9 tubercles; suture
^2 1
slightly ridged on declivity; elytra 1.8 to 1.85 x as long as wide; 1.8 to
^3 1
2.6 mm: Taphrorychus alni PFEFFER *
^5 1
suture on elytral declivity with small tubercles *127C*
#127C F199.A12 backto #127B
^17 1
suture ridged, on elytral declivity with 10 or 11 small tubercles; 1st and
^18 1
2nd striae indistinct; interstriae with fine, irregular punctures; male
^19 1
declivity with fine punctures (fig. 199); body dark brown, shiny, with sparse,
^20 1
long, white, filiform setae; antennae and legs yellowish brown; 1.6 to 2.3 mm:
^21 1
Taphrorychus bicolor HERBST *
^24 1
elytral declivity punctate or wrinkled, with slight furrow adjacent suture;
^25 1
elytra 2 x as long as wide, with striae; interstriae with fine row of punctures;
^26 1
pronotum longer than wide, anterior 1/2 pronotum arched; male frons with
^27 1
crown of dense, long, filiform setae; 1.3 to 1.8 mm:
^28 1
Taphrorychus ramicola REITTER *
#137A F211-212.A12 F213-214.A12 backto #31C
^1 1
Pityokteines: 1.6 to 3 mm; funicle 5-segmented; club nearly round, anterior
^2 1
with 3 bowed sutures (fig. 211); pronotum not constricted, anterior 1/2 with
^3 1
fine tubercles, posterior 1/2 punctate; anterior elytra slightly constricted;
^4 1
male elytra declivity with 3 lateral spines; female declivity with 3 small
^5 1
lateral tubercles; male frons with short, filiform setae; female frons and
^6 1
anterior pronotal margin with dense, long, blond, filiform setae; 2 tubercles
^7 1
between 3rd spine and posterior elytral margin; polygamous; host: conifers.
^9 1
declivital spines horizontally oriented (fig. 212); 2nd spine nearly as long
^10 1
as 3rd spine, distally hooked, and medially oriented; 3rd spine conical,
^11 1
pointed; 2 conical tubercles between 2nd and 3rd spine; female filiform setae
^12 1
of anterior pronotal margin 2 x as long as setae of frons (figs. 213, 214);
^13 1
body black to brown; 2 to 2.8 mm: Pityokteines spinidens REITTER *
^15 1
1st elytral spine conical, dorsally or nearly dorsally oriented *139A*
#139A F215-218.A12 backto #137A
^1 1
1st spine oriented perpendicular to elytra
^2 1
(fig. 215); 2nd spine nearly as long as
^3 1
3rd spine, distally hooked, horizontally
^4 1
oriented; 3rd spine conical, pointed, close
^5 1
to posterior elytral margin; 2 conical
^6 1
tubercles between 2nd and 3rd spines; female
^7 1
filiform setae of frons as long as filiform
^8 1
setae of anterior pronotal margin (figs. 216,
^9 1
217); female frons with medial tubercle
^10 1
or indentation; body black/brown; 2.5 to 3 mm:
^11 1
Pityokteines curvidens GERMAR *
^18 1
1st elytral spine oriented dorsal posteriorly
^19 1
(fig. 218); 2nd spine nearly as long as
^20 1
3rd spine, distally hooked, horizontally
^21 1
oriented; 3rd spine conical; 1 conical
^22 1
tubercle between 2nd and 3rd spines;
^23 1
female filiform setae of frons as long
^24 1
as filiform setae of anterior pronotal
^25 1
margin (figs. 219, 220); female frons
^26 1
with medial longitudinal ridge; body
^27 1
black to brown; 1.7 to 2.5 mm:
^28 1
Pityokteines vorontsovi JACOBASON *
#147A F234.A12 F232-233.A12 backto #31D
^1 1
Ips: 2.2 to 8.2 mm: funicle 5-segmented; 1st interstria before elytral
^2 1
declivity with row of fine granules; declivity steep; spines of lateral
^3 1
declivital margin longer in male than in female; medial tubercle of frons larger
^4 1
in male than in female; male filiform setae on medial posterior margin of 7th
^5 1
abdominal sternite shorter than in female (figs. 232, 233); polygamous; conifers
^7 1
1st spine on elytral declivity distally thickened (fig. 234) *147B*
^9 1
1st spine on elytral declivity distally not thickened *151A*
#147B F234.A12 backto #147A
^1 1
lateral margin of elytral declivity with 6 spines (fig. 234); 4th spine longest
^2 1
and distally thickened; 3rd and 4th spines occurring together on posterior
^3 1
protrusion of lateral margin of elytral declivity; body brown; 5.5 to 8.2 mm:
^4 1
Ips sexdentatus BOERNER *
^6 1
lateral margin of elytral declivity with 4 spines *147C*
#147C F235.A12 backto #147B
^1 1
2nd and 3rd spines of lateral elytral declivity separated at base *149A*
^3 1
2nd and 3rd spines together on posterior protrusion of lateral margin of
^4 1
elytral declivity; male 3rd spine thickened, pointed in female (fig. 235);
^5 1
elytra slightly longer than pronotum; declivity shiny, finely punctured; 3.2
^6 1
to 4 mm: Ips duplicatus SAHLBERG *
#149A F236-237.A12 backto #147C
^2 2
declivity dull, indistinctly punctured, without filiform setae (fig. 236);
^3 2
sutures of antennal club bowed (fig. 237); body posteriorly slightly wider;
^4 2
body dark brown; female anterior pronotum with more dense setae than male;
^5 2
4.2 to 5.5 mm: Ips typographus L. *
^7 2
elytral declivity shiny, with distinct punctures *149B*
#149B F238-240.A12 backto #149A
^1 1
sutures of antennal club straight (fig. 238); suture on elytral declivity
^2 1
without adjacent row of filiform setae (fig. 239);
^3 1
body dark brown; 3.5 to 4.8 mm:
^4 1
Ips amitinus EICHHOFF *
^6 1
sutures of antennal club bowed; row of long,
^7 1
blond, filiform setae adjacent suture on
^8 1
elytral declivity (fig. 240); body cylindrical;
^9 1
declivity perpendicularly ventral to 2nd spine;
^10 1
body dark brown; 4.9 to 6 mm: Ips cembrae HEER *
#151A F241-242.A12 F243-244.A12 backto #147A
^19 1
Lateral margin of elytral declivity with 3 spines; 3rd spine distally bifurcate
^20 1
in male (fig. 241), uniformly pointed in female (fig. 242); elytral slightly
^21 1
longer than pronotum; declivity shiny, with punctures denser adjacent suture;
^22 1
posterior elytral margin wider than anteriorly; body dark brown; elytra
^23 1
reddish brown; antennae and legs yellow-red; 2.2 to 3.5 mm:
^24 1
Ips acuminatus GYLLENHAL *
^25 5
Lateral margin of elytral declivity with 4 pointed spines; male 2nd spine
^26 1
slightly distally thickened (fig. 243); 2nd and 3rd spines together on
^27 1
posterior protrusion of lateral margin of elytral declivity (fig. 244);
^28 1
declivity nearly round, shiny, densely punctured; margin adjacent elytral
^29 1
suture smooth on declivity; body dark brown; elytra and legs brown; antennae
^30 1
light brown; 3 to 3.4 mm: Ips mannsfeldi WACHTL *
#141A F221-222.A12 backto #31D
^1 1
Orthotomicus: 1.6 to 5 mm; body cylindrical; funicle 5-segmented; frons with
^2 1
sparse short, filiform setae; elytra with rough striae; 1st interstria adjacent
^3 1
declivity without row of granules; declivity nearly perpendicular; tips of
^4 1
posterior elytral margin rounded, with fine indentations, not bordered; male
^5 1
elytral declivity with conical spines; female declivity with granules;
^6 1
polygamous; host: conifers.
^7 5
2nd elytral spine broad, almost flat, male with small, inclined, pointed tip
^8 1
(fig. 221); female without pointed tip; male with 2 tubercles between 2nd and
^9 1
3rd spines; female suture on declivity with row of granules; club broad, with
^10 1
bowed sutures (fig. 222); male frons with medial tubercle, larger than in
^11 1
female; body dark brown, shiny, with brown to grey filiform setae; antennae
^12 1
and legs reddish brown; 3.3 to 5 mm: Orthotomicus longicollis GYLLENHAL *
^14 1
2nd elytral spine not broad; declivity with more than 3 spines or tubercles
^15 1
*141B*
#141B backto #141A
^1 1
1 or 2 indentations between 3rd conical spine and posterior elytral margin;
^2 1
1 tubercle between 2nd and 3rd spines *143A*
^4 1
lateral declivital margin between 3rd conical spine and posterior elytral margin
^5 1
without indentations; 2 tubercles between 2nd and 3rd conical spines *145A*
#143A F223-224.A12 backto #141B
^1 1
antennal club round, with 2 slightly bowed sutures (fig. 223); 3 conical spines
^2 1
of lateral elytral declivity equidistant (fig. 224); male's 2nd spine broad,
^3 1
nearly square; lateral margin of 2nd spine horizontal, tip perpendicular;
^4 1
1st spine on 2nd interstria before declivity; 3rd interstria ends with granule
^5 1
before declivity; body black to reddish brown, shiny, with grey filiform
^6 1
setae; elytra reddish brown; 2.7 to 3.5 mm: Orthotomicus erosus WOLLASTON *
^8 1
1st and 2nd spines of elytral declivity close together *143B*
#143B F225-226.A12 backto #143A
^18 1
1 tubercle between 2nd and 3rd conical spines of elytral declivity (fig. 225);
^19 1
1st spine on 2nd interstria before declivity; 3rd interstria ends with tubercle
^20 1
before declivity; tip of 2nd conical spine extends medially as far as granule
^21 1
of 3rd interstria; striae indistinct; sutures of club bowed (fig. 226); body
^22 1
dark, reddish-brown; 3 to 3.8 mm: Orthotomicus proximus EICHHOFF *
^25 1
1 small tubercle between 2nd and 3rd conical spines of elytral declivity; 3rd
^26 1
conical spine slightly medial to lateral declivital margin; elytral suture
^27 1
ridged on declivity; pronotum longer than wide; body brown, with light brown,
^28 1
filiform setae; antennae and legs light brown; 1.6 to 2 mm:
^29 1
Orthotomicus starki SPESSIVTSEFF *
#145A F227-230.A12 F228-231.A12 backto #141B
^1 25
sutures of club straight (fig. 227); elytral interstriae
^2 25
with fine rows of punctures; declivity round, distinctly
^3 25
bordered (fig. 228); distance between contralateral 1st
^4 25
spines similar to distance between ipsilateral 1st and
^5 25
2nd spines (fig. 229); body reddish brown; 2.5 to 4 mm:
^6 25
Orthotomicus laricis FABRICIUS *
^18 25
sutures of antennal club bowed (fig. 230); elytral
^19 25
interstriae wrinkled, with punctures; declivity oval,
^20 25
indistinctly bordered (fig. 231); distance between
^21 25
contralateral 1st spines longer than between
^22 25
ipsilateral 1st and 2nd spines; body dark brown;
^23 25
elytra brown; 2.3 to 3.5 mm:
^24 25
Orthotomicus suturalis GYLLENHAL *
#87A backto #27C
^1 1
Crypturgus: 1 to 1.5 mm: antennae and legs light brown; funicle 2-segmented;
^2 1
club broad, irregularly oval; club with indistinct suture; pronotum longer than
^3 1
wide; elytral declivity round; sexual dimorphism: length and form of pronotum;
^4 1
monogamous; host: conifers.
^6 1
elytra shiny; interstriae wiht sparse, filiform setae; punctures of striae
^7 1
round *87B*
^9 1
elytra dull; interstriae narrow, wrinkled, with fine, dense, filiform setae and
^10 1
rows of punctures; punctures of striae oval; pronotum densely punctured *89A*
#87B F159-160.A12 backto #87A
^13 1
elytra with rough striae (fig. 159); interstriae with rows of white to light
^14 1
brown, short, thick, filiform setae; pronotum fine, sparsely punctured; body
^15 1
black to brown; 1.2 to 1.3 mm: Crypturgus hispidulus THOMSON *
^17 1
elytra 2 x as long as wide, with striae (fig. 160); interstriae with rows of
^18 1
short, sparse, filiform setae; reticular pronotum longer than wide, with fine
^19 1
punctures; body black to brown; 1 to 1.2 mm: Crypturgus pusillus GYLLENHAL *
#89A F161.A12 backto #87A
^1 1
reticular pronotum fine, sparsely punctured, dull (fig. 161); sexual dimorphism:
^2 1
male pronotum longer than wide; elytra with striae; filiform setae of
^3 1
interstriae as long as those of striae; body black to brown; 1.1 to 1.5 mm:
^4 1
Crypturgus cinereus HERBST *
^6 1
pronotum shiny, longer than wide, with dense, rough punctures; elytra with
^7 1
rough, granulate striae *89B*
#89B F162-163.A12 backto #89A
^12 1
pronotum not anteriorly constricted, with dense, fine punctures; sexual
^13 1
dimorphism: female elytral declivity with dense, filiform setae (fig. 162);
^14 1
body black to brown; 1.1 to 1.5 mm: Crypturgus mediterraneus EICHHOFF *
^15 3
punctate pronotum anteriorly constricted (fig. 163); sexual dimorphism: female
^16 1
pronotum short, wide; male pronotum shorter and narrower; male elytral declivity
^17 1
adjacent suture with round, dull area, without filiform setae; female declivity
^18 1
with dense, filiform setae; body dark brown; antennae and legs light brown;
^19 1
1 to 1.4 mm: Crypturgus numidicus FERRARI *


